This is Mazon Monday post #94. What's your favorite Mazon Creek fossil? Tell us at email:[email protected].
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By Falconaumanni - Own work, CC BY-SA 3.0, https://commons.wikimedia.org/w/index.php?curid=56797814
For this week, we are looking at Calamites, which is a genus of extinct arborescent (tree-like) horsetails. They are sphenopsids. Modern horsetails, genus Equistem, are fairly closely related. Some of these Carboniferous plants could grow to heights of more than 100 feet (30+ meters). Additionally, they were a key component of the forest understory and make a large part of Pennsylvanian coal deposits. There are a few species of Calamites known in the Mazon Creek biota - including Calamites cistii, . Also, note that while Calamites refers to the stem of sphenopsids, Annularia (see Mazon Monday #60), Asterophyllites, and Equisetites are other parts of the same group of plants. George Langford has a great picture that details the individual parts.
This way of describing plants with different species for each part is called Form Genera. It's due to the often fragmentary fossil preservation of plants. A "Key" specimen has multiple associated plant parts and is used to tie the separate parts together. The separate species names for plant parts allows paleontologists to discuss the various fossils that are found. There is a good explanation in the concept in the "Keys to Identify Pennsylvanian Fossil Plants of the Mazon Creek Area".
THE CONCEPT OF FORM-GENERA IN IDENTIFYING FOSSIL PLANTS
Biologists and paleobiologists who study and classify organisms often find biological structures that cannot be confidently assigned to any known organismic species. Such structures -- mostly parts, life-stages, or sedimentary traces of organisms contain considerable biological information and provide insights into the morphology, life-style, and behavior of past (and present) species. For example, for many present-day fungi, both the asexual and sexual stages of the life-cycle is known, but each stage cannot be causally linked to the other. These stages may have separate, taxonomic names, yet belong to the same species. Similarly, trace fossils distinctively-shaped impressions and burrows indicating particular behavioral habits are given separate taxonomic names, and are not assuredly linked to any known fossil species.
And so it is with fossil plants, in which particular organs are preserved in isolation from other organs belonging to the same plant species. Thus plant leaves, roots, stems, fruits, seeds, and even pollen or spores, have separate taxonomic names. These names are termed "form genera". Several form-genera, each designating a specific part of the plant, can refer to a single fossil plant species, but it is often impossible to know which leaf-genus is associated with which seed-genus or pollen-genus, and so on.
Furthermore, in some cases it is clear that structures assigned to one form were actually produced by two or more different kinds of plants, e.g., Lepidophylloides produced by both Sigillaria and Lepidodendron. Therefore, even when we can demonstrate, or hypothesize, that two or more form-genera are actually part of a single plant species, we cannot be certain that all fragmentary specimens of the genera are correctly treated in this way. For these reasons, form-genera are an indispensable, if slightly confusing, tool in identifying fossil plants.
For this post, we are looking at the stem of Calamites, of which Calamites cistii is the most common. Here is its entry in "A Comprehensive Guide to the Fossil Flora of Mazon Creek".
Calamites cistii Brongniart, 1828
1828. Calamites cistii Brongniart: p. 129, pl. 20
1879-80. Calamites cistii Brongniart; Lesquereux: p.
1899. Calamites cistii Brongniart; White: p. 149 27, pl. 1, fig. 6
1958. Calamites cisti Brongniart; Langford: p. 31, fig. 22
1969. Calamites cisti Brongniart; Crookall: p. 639, pl.
1969. Calamites cistii Brongniart; Darrah: p. 171, 123, fig. 1
1979. Calamites cisti Brongniart; Janssen: p. 80, fig. 61
DESCRIPTION: The nodes are not contracted and are regularly spaced along the stem. The internodes are longer than they are wide. The ribs are thin, straight, half-round at their ends, and separated by striate furrows. The vascular canal scars, if present, are small, round, and indistinct. As with all calamite stems, only fossils formed as pith casts will have vascular canal scars. All figures shown here are of outer surfaces of stems.
REMARKS: Calamites in general are common, and Calamites cistii is the most frequently found species of this group in the Mazon Creek flora. It differs from Calamites suckowi in its more consistent internode length and narrower ribs.
Specimens
From George's Basement (George Langford)