This is Mazon Monday post #195. What's your favorite Mazon Creek fossil? Tell us at email:[email protected].
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Figure 20.4. Vetacapsula cooperi, modified from McGhee and Richardson (1982). Scale bar 10 mm.
Vetacapsula cooperi is a very rare shark egg case from the Pennsylvanian Period. It was originally described by Robert Crookall (1890-1981) in 1928. In the late 1920's, Crookall took a deep look at Carboniferous shark egg cases, then classified as enigmatic flower-buds or seed-vessels, and provided a comprehensive overview of all the species of Palaeoxyris, Fayolia, and Vetacapsula known at the time. See page 80 of "Elasmobranch egg capsules Palaeoxyris, Fayolia, and Vetacapsula as subject of paleontological research - and annotated bibliography".
Between 1928 and 1932 R. Crookall provided a complete and comprehensive overview of all species of Palaeoxyris, Fayolia and Vetacapsula known at the time (Crookall, 1928a, 1928b, 1930, 1932). Furthermore, at the same time he revised and erected several new species (P. warei, P. pringlei, P. edwardsi, P. duni, P. bohemica, F. cambriensis, F. eltringhami, F. besti, F. warei, F. interstrialis, V. kidstoni, V. moyseyi, and V. hemingwayi). In Crookall (1932) the nature and affinities of these fossils were discussed in detail. First he listed 12 different groups to which Palaeoxyris and related genera Fayolia and Vetacapsula had been referred or with which they had been compared. He demonstrated that all remains invariably occur in fresh- or brackish water deposits; the capsules were attached to plants by their pointed beaks, and never showed cellular issue, which precludes them from being plant remains. Crookall came to the conclusion that they were of animal origin and had to be accepted as egg capsules of elasmobranchs. Furthermore, Crookall cited D. Watson’s remark about the shark family Hybodontidae, whose stratigraphical range covers exactly that of Palaeoxyris. With regard to Fayolia and Vetacapsula, other possible forms (Ichthyotomi, Petalodonttidae, Cochliodontidae, Cladontidae) were mentioned; their geological range, however, provided no assistance.
It was unknown from Mazon Creek until it was discovered in the back of a drawer at the Field Museum of Natural History in Chicago. The discovery prompted George McGhee and Eugene Richardson to write "First occurrence of the problematic fossil Vetacapsula in North America". The paper was published in 1982 in the Journal of Paleontology.
V. cooperi can be found on page 163 of Jack Wittry's "The Fossil Fauna of Mazon Creek".
Vetacapsula cooperi (Mackie) Crookall, 1928
Vetacapsula was unknown in North America until 1982 when an example was found hidden in a drawer of The Field Museum. As with Palaeoxyris (see page 159), it too was a problematic fossil, but now its affinities are agreed upon. They are considered to be the egg capsules for an unknown type of chondrichthyan shark.
V. cooperi has a heavily inflated, spindle-shaped body. Its surface is marked by straight, pleat-like ribs spaced about 1 mm apart that run from the pedicle to the distal end. A strong median ridge running more or less the length of the body has been reported, but is not always seen.
Specimens
From "The Fossil Fauna of Mazon Creek"
From ESCONI member Bob Masek
Vetacapsula sp. from Crock Hey. Similar species.